Who shot Putin-critic Boris Nemtsov – and why?

Russia's top investigative body said Saturday it is looking into several possible motives for the killing of prominent opposition figure Boris Nemtsov, including an attempt to destabilize the state, Islamic extremism, the Ukraine conflict and his personal life.

A statement from the body, the Investigative Committee, did not address the possibility seen as likely by many of Nemtsov's supporters — that he was killed for being one of President Vladimir Putin's most adamant and visible critics.

The 55-year-old Nemtsov was gunned down Friday near midnight as he walked on a bridge near the Kremlin with a female companion. The killing came just a few hours after a radio interview in which he denounced Putin's "mad, aggressive" policies and the day before he was to help lead a rally protesting Russia's actions in the Ukraine crisis and the economic crisis at home.

After his death, organizers canceled the rally and instead called for a demonstration to mourn him on Sunday in central Moscow. The city gave quick approval for that gathering, in contrast to its usual slow and grudging permission for opposition rallies.

The Investigative Committee said it was looking into whether Nemtsov had been killed as a "sacrificial victim for those who do not shun any method for achieving their political goals," a suggestion echoing the comments by Putin's spokesman and other Russian politicians that the attack was a "provocation" against the state.

It also said it was considering whether there was "personal enmity" toward him in his domestic life. State-controlled TV and Kremlin-friendly media outlets on Saturday gave considerable attention to Nemtsov's companion, identifying her as a Ukrainian model 30 years his junior and showing photos of her in alluring poses. The Investigative Committee said the pair were headed for Nemtsov's apartment.

The statement also said it was investigating whether the killing was connected to the Ukraine conflict, where Russia-backed separatists have been fighting Ukrainian forces since last April, or whether there was a connection to Islamic extremism.

Nemtsov had been one of Putin's most visible critics and his death hit other members of the opposition hard. The mourning march on Sunday could serve to galvanize the beleaguered and marginalized opposition, or it could prove to be a brief catharsis after which emotions dissipate.

Through the morning, hundreds of people came to the site of Nemtsov's death to lay flowers.

Nemtsov was working on a report presenting evidence that he believed proved Russia's direct involvement in the separatist rebellion that has raged in eastern Ukraine since April. Moscow denies backing the rebels with troops and sophisticated weapons.

Putin ordered Russia's top law enforcement chiefs to personally oversee the investigation of Nemtsov's killing.

"Putin noted that this cruel murder has all the makings of a contract hit and is extremely provocative," presidential spokesman Dmitry Peskov said in remarks carried by Russian news agencies.

President Barack Obama said the Russian people "lost lost one of the most dedicated and eloquent defenders of their rights."

German Chancellor Angela Merkel praised Nemtsov's courage in criticizing Kremlin policies, and urged Putin to insure that the killers are brought to justice, her spokesman Steffen Seibert said.

Former Soviet leader Mikhail Gorbachev echoed the suggestion that the killing was a provocation. "It's an attempt to push the situation into complications, maybe even to destabilizing the situation in the country," he was quoted as saying by the Interfax news agency.

Communist Party leader Gennady Zyuganov agreed. "It's a provocation; for big fires, sacrificial figures are necessary," Interfax quoted him as saying.

Nemtsov frequently assailed the government's inefficiency, rampant corruption and Ukraine policy.

In an interview with the Sobesednik newspaper, Nemtsov said earlier this month that his 86-year old mother was afraid that Putin could have him killed. Asked if he had such fears himself, he responded: "If I were afraid I wouldn't have led an opposition party."

Speaking on radio just a few hours before his death, he accused Putin of plunging Russia into crisis by his "mad, aggressive and deadly policy of war against Ukraine."

Kasyanov, the former prime minister, said he was shocked.

"In the 21st century, a leader of the opposition is being demonstratively shot just outside the walls of the Kremlin!".

"This is a monstrous tragedy and a loss for us all," Alexei Navalny, Russia's most prominent opposition figure, said on his Facebook page. He is currently on a 15-day jail sentence for handing out leaflets without authorization.

"The country is rolling into the abyss," Kasyanov, the former prime minister, told reporters as Nemtsov's body, placed in a plastic bag, was removed on a rainy and cold night, as the Kremlin bells chimed nearby.

Nemtsov served as a regional governor and then a deputy prime minister in the 1990s and once was seen as a possible successor to Boris Yeltsin, Russia's first elected president.

Nemtsov was widely liked for his good humor, larger-than-life character and quick wit, but he and other top opposition figures long have been purged from state television and steadily marginalized by the Kremlin.

Associated Press writers Vladimir Isachenkov and Laura Mills in Moscow and Jake Pearson in New York contributed to this report.